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Sense organs 367

1

 

 

 

 

 

 

1

 

 

 

 

 

2

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

3

 

 

20

16

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

3

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

12

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

9

11

8

 

17

 

4

 

10

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

18

5

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

4

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

6

 

 

Palpebral fissure

 

 

 

 

Tarsal plates and ligaments

 

 

 

 

 

 

 

 

 

A

 

 

 

 

 

 

 

 

B

7

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

8

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

9

 

 

 

22

 

 

 

 

 

 

6

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

10

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

11

 

 

 

 

 

 

 

 

 

 

 

 

 

16

21

5

 

7

 

14

13

8

 

21

15

13

7

 

 

17

24

5

 

23

E Eyelids, sagittal section

 

 

 

 

 

 

 

C

Epicanthus (epicanthic fold)

 

 

12

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

13

 

 

 

 

 

 

 

 

 

 

 

 

 

 

19

 

 

 

18

14

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

15

 

 

 

 

 

 

 

16

D

Orbicular muscle of eye

17

 

from behind

 

 

18

19

20

 

 

 

15

 

 

 

 

 

 

 

 

 

 

 

 

 

21

 

 

 

 

 

25

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

22

 

 

 

 

 

 

 

 

 

26

 

 

 

 

 

 

23

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

24

 

Inner (nasal) canthus of eye

 

 

F

 

25

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

368

Sense organs

 

 

 

 

 

 

 

 

 

1

 

Bulbar conjunctiva. Tunica conjunctiva bul-

 

15

Rivus lacrimalis. Pathway that conducts tears

1

 

 

 

 

 

 

baris. Part of conjunctiva covering the eyeball. It

 

 

from the excretory ducts to the lacrimal lake. It

 

 

 

 

 

consists of stratified, nonkeratinized squamous

 

 

lies within the conjunctival sac between the

2

 

 

 

 

epithelium with only a few goblet cells and a

 

 

closed eyelids and the eyeball.

 

 

 

 

 

loose, cell-poor lamina propria permeated with

 

16

Lacrimal lake. Lacus lacrimalis. Space in the me-

 

 

 

 

 

 

3

 

 

 

 

elastic fibers. A

 

 

 

dial angle of

the eye

around the lacrimal

 

 

 

 

 

 

 

 

2

 

Palpebral conjunctiva. Tunica conjunctiva pal-

 

 

caruncle. B C

 

 

 

 

 

 

 

 

 

 

 

 

 

pebralis. The portion of the conjunctiva covering

 

17 Papilla lacrimalis. Small cone-shaped elevation

4

 

 

 

 

 

 

 

 

 

the posterior surface of the eyelid. It consists of

 

 

 

 

 

 

 

medial to the inner edge of both the upper and

 

 

 

 

 

two or more layers of columnar epithelium with

 

 

lower eyelids. Each apex houses an opening or

 

 

 

 

 

 

 

5

 

 

 

 

goblet cells and a loose, vascularized lamina

 

 

lacrimal punctum. C

 

 

 

 

 

propria. A

 

 

 

 

 

 

 

 

 

 

 

18 Lacrimal punctum. Punctum lacrimale. Small

 

3

 

Superior fornix of

cunjunctiva. Fornix con-

 

6

 

 

 

opening marking the beginning of the lacrimal

 

 

 

 

junctivae superior. Reflected fold of conjunctiva

 

 

fluid drainage system. C

 

 

 

 

 

 

 

 

 

 

 

 

 

 

extending from the eyeball (bulbar) to the upper

 

19

Lacrimal

canaliculus. Canaliculus lacrimalis.

7

 

 

 

 

eyelid (palpebral). A

 

 

 

 

 

 

 

 

 

Small canal, up to 1 cm long, from each lacrimal

4

 

Inferior fornix of conjunctiva. Fornix conjunc-

 

 

 

 

 

 

punctum to the lacrimal sac. C

8

 

 

 

 

tivae inferior. Reflected fold of conjunctiva from

 

20

Ampulla

of

lacrimal

canaliculus. Ampulla

 

 

 

 

 

the eyeball (bulbar)

on to the lower eyelid

 

 

canaliculi lacrimalis. Slight enlargement at the

 

 

 

 

 

(palpebral). A

 

 

 

9

 

 

 

 

 

 

 

bend of the lacrimal canaliculus. C

5

Conjunctival sac. Saccus conjunctivalis. Space

 

 

 

21 Lacrimal sac. Saccus lacrimalis. It is located in

 

 

 

 

 

 

 

between palpebral and bulbar conjunctivae. Its

 

10

 

 

 

 

 

 

the lacrimal fossa and is about 1.5 cm long and

 

 

 

 

upper and lower ends form the superior and in-

 

 

 

 

 

 

 

 

about 0.5 cm wide. It descends directly into the

 

 

 

 

 

ferior fornices of the conjunctiva. A

 

 

nasolacrimal duct. C

 

11

6

 

Ciliary glands (of Moll). Glandulae ciliares

 

 

 

 

 

22

Fornix of lacrimal sac. Fornix sacci lacrimalis.

 

 

 

 

 

[[Molli]]. Apocrine glands on the lid margin.

 

 

Dome-shaped upper margin of the lacrimal sac.

 

 

 

 

 

 

 

12

 

 

 

 

They open either into the hair follicles of the

 

 

C

 

 

 

 

 

 

 

eyelashes or at the lid margin. A

 

23 Nasolacrimal duct. Ductus nasolacrimalis. Duct

 

7

 

Sebaceous glands (of Zeiss). Glandulae se-

 

13

 

 

 

that is directly continuous with the larcrimal sac

 

 

 

 

baceae [[Zeiss]]. Small sebaceous glands with

 

 

and about 1.2−2.4 cm in length. It passes

 

 

 

 

 

openings into the hair follicles of the eyelashes.

 

 

through the nasolacrimal canal and opens into

14

 

 

 

 

A

 

 

 

the inferior nasal meatus. Its flattened lumen is

 

8

 

Conjunctival glands. Glanduale conjunctivales.

 

 

lined by a mucosa containing two or more layers

 

 

 

 

 

 

 

 

of columnar epithelium bearing cilia at some

15

 

 

 

 

Follicular aggregations of lymphocytes at the

 

 

 

 

 

 

 

 

sites. C

 

 

 

 

 

 

 

medial angle of the eye.

 

 

 

 

 

 

9

 

Lacrimal apparatus. Apparatus lacrimalis. The

 

24

Lacrimal fold. Plica lacrimalis. Mucosal fold at

16

 

 

 

 

 

the opening of the nasolacrimal duct. It is lo-

 

 

 

 

system of structures that lubricate the cornea

 

 

 

 

 

 

 

 

 

cated in the inferior nasal meatus about 3−

 

 

 

 

 

and conjunctiva. B

 

 

 

17

 

 

 

 

 

 

 

3.5 cm posterior to the external naris. C

10

Lacrimal gland. Glandula lacrimalis. Gland lo-

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

cated above the lateral angle of the eyelids; it is

 

 

 

 

 

 

18

 

 

 

 

separated into an upper and lower portion by

 

 

 

 

 

 

 

 

 

 

 

the tendon of levator palpebrae muscle. Its ex-

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

cretory ducts open laterally into the superior

 

 

 

 

 

 

19

 

 

 

 

 

 

 

 

 

 

 

 

 

 

fornix of the conjunctiva. B

 

 

 

 

 

 

 

11

 

Orbital part. Pars orbitalis. Larger portion of

 

 

 

 

 

 

20

 

 

 

 

 

 

 

 

 

 

 

lacrimal gland located above the tendon of the

 

 

 

 

 

 

 

 

 

 

 

levator palpebrae muscle. B

 

 

 

 

 

 

21

12

 

Palpebral part. Pars palpebralis. Smaller por-

 

 

 

 

 

 

 

 

 

 

 

tion of lacrimal gland located below the tendon

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

22

 

 

 

 

of the levator palpebrae muscle. B

 

 

 

 

 

 

 

13

Excretory ducts of

lacrimal gland. Ductuli

 

 

 

 

 

 

 

 

 

 

 

 

 

 

23

 

 

 

 

exretorii [[glandulae

lacrimalis]]. 6−14 ducts

 

 

 

 

 

 

 

 

 

 

opening into the superior fornix of the conjunc-

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

24

 

 

 

 

tiva. B

 

 

 

 

 

 

 

 

14

 

Accessory lacrimal glands. [Gll. lacrimales ac-

 

 

 

 

 

 

 

 

 

 

 

cessoriae]. Additional smaller lacrimal glands

 

 

 

 

 

 

25

 

 

 

 

found scattered especially in the vicinity of the

 

 

 

 

 

 

 

 

 

 

 

superior conjunctival fornix. A

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

Sense organs

369

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

5

 

 

 

3

 

 

 

 

 

 

 

 

 

 

 

1

 

 

1

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

2

 

 

 

 

 

 

 

 

 

 

2

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

11

12

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

3

 

 

 

 

 

 

 

 

14

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

4

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

5

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

13

 

 

 

 

 

 

 

 

 

6

7

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

16

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

Lacrimal gland

 

 

7

 

 

 

 

 

 

 

 

 

 

 

 

 

 

B

 

 

 

6

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

8

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

9

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

10

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

11

2

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

5

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

20

 

 

 

 

 

 

12

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

17

19

 

 

 

 

 

 

 

 

4

 

 

 

13

 

 

A

Eyelids, sagittal section

 

 

 

 

18

 

 

 

 

 

 

22

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

16

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

14

 

 

 

 

 

 

 

 

 

 

 

 

17

18

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

19

21

 

 

15

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

20

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

16

17

18

23

19

20

24

21

22

C Lacrimal system

23

24

25

 

370

Sense organs

 

 

 

 

 

 

 

 

 

 

 

 

 

1

VESTIBULOCOCHLEAR ORGAN. Organum vesti-

16

Basal membrane of semicircular duct. Mem-

1

 

 

bulocochleare. Sensory apparatus housed in the

 

brana basalis ductus semicircularis. Appears

 

 

 

temporal bone for the perception of sound,

 

upon light microscopy as a homogeneous basal

2

 

 

equilibrium and positional changes.

 

membrane situated directly below the

 

2

INTERNAL EAR. Auris interna. Part of the vesti-

 

epithelium. C

 

 

 

 

 

 

 

17

 

 

 

 

 

 

 

 

3

 

 

bulocochlear organ residing in the petrous tem-

[[Epithelium

of

semicircular

duct]].

 

 

poral bone.

 

 

 

[[Epithelium

ductus

semicircularis]]. Simple

 

 

 

 

 

 

 

3

MEMBRANOUS LABYRINTH. Labyrinthus mem-

 

epithelium lining the inner aspect of the mem-

4

 

branous semicircular duct. The cells are flat and

 

 

branaceus. Complicated system of ducts and di-

 

 

 

 

latations within the bony labyrinth, which con-

 

become cuboidal on their concave side. C

 

 

 

 

 

 

5

 

 

tains sensory epithelium and is suspended by

18

Membranous ampullae. Ampullae mem-

 

 

 

connective tissue. A

 

 

 

branaceae. Dilatations of the semicircular ducts

 

4

Endolymph. Fluid contained within the mem-

 

in the vicinity of the utricle.

 

 

 

6

 

 

 

 

19

Anterior

membranous

ampulla.

Ampulla

 

 

branous labyrinth.

 

 

 

 

 

 

 

 

5

Perilymph. Fluid occupying the osseous laby-

 

membranacea anterior. Dilatation at the end of

7

 

anterior (superior) semicircular duct located

 

 

rinth and surrounding the membranous laby-

 

 

 

 

rinth.

 

 

 

near the lateral membranous ampulla. A

 

 

 

 

 

 

 

 

8

 

 

 

 

20

Posterior

membranous

ampulla. Ampulla

6

Vestibular labyrinth. Labyrinthus vestibularis.

 

 

 

Portion of membranous labyrinth constituting

 

membranacea posterior. Dilatation at the end of

 

 

 

 

the posterior semicircular duct located distal to

9

 

 

the organ of equilibrium. It includes the semi-

 

 

 

 

the other two membranous ampullae. A

 

 

 

circular ducts.

 

 

 

 

 

 

 

 

 

21

Lateral

membranous

ampulla.

Ampulla

 

7

Endolymphatic duct. Ductus endolymphaticus

 

10

 

membranacea lateralis. Ampulla of the lateral

 

 

[[aquaeductus vestibuli]]. Slender duct arising

 

 

 

 

 

semicircular duct located proximal to the ante-

 

 

 

from the utriculosaccular duct and passing

 

 

 

 

 

rior membranous ampulla. A

 

 

 

11

 

 

through the osseous aqueduct of the vestibule to

 

 

 

 

 

 

22

Sulcus ampullaris. Indentation below the am-

 

 

 

terminate as the endolymphatic sac. A

 

8

Endolymphatic sac. Saccus endolymphaticus.

 

pullary crest bearing branches from the ampul-

12

 

 

lar nerve for innervation of the ampullary crest.

 

 

Blind sac of endolymphatic duct located be-

 

 

 

 

 

B

 

 

 

 

 

 

 

 

 

 

tween two dural layers at the posterior wall of

 

 

 

 

 

 

 

 

13

 

 

23

Ampullary crest. Crista ampullaris. Crescent-

 

 

the petrous temporal. A

 

 

9

Utriculosaccular duct. Ductus

utriculosaccu-

 

shaped ridge

projecting

into

the

ampullary

 

 

14

 

 

laris. Slender duct between the saccule and

 

space. It is covered by sensory epithelium and

 

 

 

has a base of nerve fibers and connective tissue.

 

 

 

utricle. It gives rise to the endolymphatic duct. A

 

B

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

15

10

Utricle. Utriculus. Sac 2.5−3.5 mm in diameter,

 

 

 

 

 

 

 

 

24

[[Neuroepithelium]].

Sensory

epithelium

of

 

 

 

serving as the base for the three semicircular

 

ampullae

consisting of supporting

cells

and

 

 

 

ducts. A

 

 

 

16

11

Ductus

semicirculares.

 

sensory cells with hairs (microvilli) projecting

Semicircular duct.

 

 

from the surface into an overlying cupula. B

 

 

 

Three membranous ducts that resemble two-

25

Cupula. Gelatinous body suspended above the

17

 

 

thirds of a circular arch, each occupying its own

 

 

 

ampullary crest as far as the roof of the ampulla

 

 

 

osseous semicircular canal oriented perpendic-

 

and penetrated by hairs of the sensory cells. B

 

 

 

 

 

 

 

ular to the others.

 

 

 

18

 

 

 

 

 

 

 

 

 

26

Membranous

crura.

Crura

membranacea.

12

Anterior (superior)

semicircular duct. Duc-

 

 

 

tus semicircularis anterior. It is oriented verti-

 

Limbs of semicircular ducts opening into the

19

 

 

 

utricle.

 

 

 

 

 

 

 

 

 

cally and somewhat perpendicular to the

 

 

 

 

 

 

 

 

 

 

27

Simple membranous crus. Crus mem-

 

 

 

petrous part of the temporal bone. A

 

 

 

20

13

Posterior semicircular duct. Ductus semi-

 

branaceum simplex. Posterior limb of lateral

 

semicircular duct opening independently into

 

 

 

circularis posterior. It is oriented somewhat ver-

 

the utricle. A

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

21

 

 

tically in a plane which runs parallel to the longi-

 

 

 

 

 

 

 

 

 

28

Ampullary membranous crura. Crura mem-

 

 

tudinal axis of the petrous part of the temporal

 

 

 

bone. A

 

 

 

branacea ampullaria. Semicircular duct seg-

22

14

Lateral semicircular duct. Ductus semicircu-

 

ments situated between the ampullae and the

 

utricle. A

 

 

 

 

 

 

 

 

 

 

laris lateralis. The most lateral, horizontally

 

 

 

 

 

 

 

 

 

 

 

29

Common

membranous

crus. Crus mem-

23

 

 

oriented semicircular duct. It may create a bulge

 

 

in the medial wall of the tympanic cavity. A

 

branaceum commune. Common limb formed by

 

 

 

 

 

15

Proper membrane of semicircular duct. Mem-

 

the anterior and posterior semicircular ducts

24

 

and opening into the utricle. A

 

 

 

 

 

brana propria ductus semicircularis. Layer

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

below the basal membrane consisting primarily

 

 

 

 

 

 

 

 

 

25of densely packed fibers which extends into the looser network of the perilymphatic space. C

Sense organs 371

 

 

12

 

 

19

 

13

 

 

28

29

14

 

 

10

21

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

27

 

 

 

25

 

 

 

9

20

7

 

 

 

 

 

 

8

A Membranous labyrinth

 

25

 

 

 

17

24

23

16

15

 

 

22

 

B Ampulla of semicircular duct

C Semicircular duct, cross section

1

2

3

4

5

6

7

8

9

10

11

12

13

14

15

16

17

18

19

20

21

22

23

24

25