8. Chan K.L., Dhaliwal S.S., Young H.S., 1979. Protein variation and systematics of three subgenera of Malayan rats (Rodentia: Muridae, genus Rattus Fischer) // Comp. Biochem. V. 648. P. 329-337.
9. Chevret P., Catzevlis F., Furano A.V., Verneau O., 1998. Comparison of the evolutionary relationships between South-East Asian rats (Rodentia: Murinae) as deduced from four molecular studies // Abstr. Euro-Amer. Mammal Congr, Santiago de Compostela, Spain. P. 18.
10. Corbet G.B., Hill J.E., 1992. The mammals of the Indomalayan region: a systematic review. Oxford: Oxford Univ. Press. 488 p
11. Davis D.J., 1964. Disc-electrophoresis. II. Method and application to human serum protein // Ann. N.Y. Acad. Sci., V. 121. P. 404-408.
12. Gadi I.K., Sharma T., 1983. Cytogenetic relationships in Rattus, Cremnomys, Millardia, Nesokia and Bandicuta (Rodentia: Muridae) // Genetica, V. 61. P. 21-40.
13. Gemmeke H., Niethammer J., 1984. Zur taxonomie der gattung Rattus (Rodentia, Muridae) // Z. Saugetierk. Bd. 49. S. 104-116.
14. Harris H., Hopkinson D.A., 1978. Handbook of enzyme electrophoresis in human genetics. Amsterdam: North-Holland. Publ. Co.
15. Likhnova O.P., Musser G.G., Kuznetsov G.V., Pavlinov I.J. 1998. Preliminary results of allozymic and cranial analyses of Niviventer (Rodentia: Muridae) from southern Vietnam // Abstr. Euro-Amer. Mammal Congr, Santiago de Compostela, Spain. P. 356.
16. Musser G.G., 1981. Results of Archbold expeditions. № 105. Notes on systematics of Indo-Malayan murid rodents, and descriptions of new genera and species from Ceylon, Sulawesi, and the Philippines // Bull. Amer. Mus. Nat. Hist. V. 168. Art. 3. P. 229-334.
17. Musser G.G., Carleton M.D., 1993. Family Muridae // Wilson D.E., Reeder D.M. (eds). Mammal species of the World. A taxonomic and geographic reference. 2d ed. Washington: Smthsonian Inst. Press. P. 501-756.
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19. Verneau O., Catzeflis F., Furano A., 1998. L1 (Line-1) petrotransposons provide unique phylogenetic characters for deciphering and dating the speciation events between species and genera in Rattus sensu lato (Rodentia: Muridae) // Abstr. Euro-Amer. Mammal Congr, Santiago de Compostela, Spain. P. 14-15.
20. Yong H.S., 1969. Karyotypes of Malayan rats (Rodentia--Muridae, genus Rattus Fischer) // Chromosoma, V. 27. P. 245-267/
Приложение
Таблица 1 Частоты аллелей полиморфных локусов в исследованных выборках крыс Вьетнама
|
Виды/Места сбора |
|||||||||||
|
Локус и |
N. tenaster |
N. fulvescens |
N. langbianis |
M. surifer |
M. moi |
B. bowersi |
L. cf. edwardsi |
R. koratensis |
R. exulans |
||
|
число экз. |
Cong Troi |
Phi Lieng |
|||||||||
|
N |
17 |
2 |
2 |
1 |
2 |
1 |
3 |
1 |
1 |
1 |
|
|
Ck-2 |
b (1.00) |
b (1.00) |
b (1.00) |
b (1.00) |
a (1.00) |
c (1.00) |
a (0.333) |
b (1.00) |
c (1.00) |
c (1.00) |
|
|
b (0.667) |
|||||||||||
|
N |
21 |
2 |
8 |
1 |
2 |
1 |
1 |
1 |
|||
|
Dia-2 |
a (1.00) |
a (1.00) |
a (1.00) |
c (1.00) |
c (1.00) |
c (1.00) |
b (1.00) |
b (1.00) |
|||
|
N |
22 |
2 |
9 |
1 |
1 |
3 |
|||||
|
Es-1 |
a (0.182) |
b (0.750) |
b (0.944) |
g (1.00) |
e (0.500) |
d (0.333) |
|||||
|
b (0.795) |
d (0.250) |
d (0.056) |
f (0.500) |
e (0.667) |
|||||||
|
c (0.023) |
|||||||||||
|
N |
21 |
2 |
8 |
1 |
|||||||
|
Es-2 |
a (0.548) |
d (0.250) |
d (0.063) |
e (1.00) |
|||||||
|
b (0.429) |
e (0.750) |
e (0.938) |
|||||||||
|
c (0.024) |
|||||||||||
|
N |
22 |
2 |
9 |
1 |
|||||||
|
Es-3 |
a (0.091) |
c (1.00) |
c (1.00) |
d (1.00) |
|||||||
|
b (0.909) |
|||||||||||
|
N |
22 |
2 |
9 |
1 |
|||||||
|
Es-4 |
a (0.144) |
d (1.00) |
d (0.722) |
c (1.00) |
|||||||
|
b (0.886) |
e (0.278) |
||||||||||
|
N |
22 |
2 |
9 |
1 |
2 |
1 |
6 |
1 |
1 |
1 |
|
|
Got-1 |
c (1.00) |
c (1.00) |
c (1.00) |
c (1.00) |
d (1.00) |
d (1.00) |
b (1.00) |
a (0.500) |
a (1.00) |
a (1.00) |
|
|
b (0.500) |
|||||||||||
|
N |
22 |
2 |
2 |
1 |
|||||||
|
Gp (почки) |
b (1.00) |
a (1.00) |
a (1.00) |
b (1.00) |
|||||||
|
N |
22 |
2 |
7 |
1 |
1 |
3 |
1 |
||||
|
Gp-1 (эритроциты) |
b (1.00) |
a (1.00) |
a (1.00) |
b (1.00) |
c (1.00) |
c (1.00) |
b (1.00) |
||||
|
N |
22 |
2 |
7 |
1 |
1 |
1 |
3 |
1 |
|||
|
Gp-2 (эритроциты) |
b (1.00) |
b (1.00) |
b (1.00) |
b (1.00) |
b (1.00) |
b (1.00) |
b (1.00) |
a (1.00) |
|||
|
N |
18 |
2 |
5 |
1 |
2 |
1 |
6 |
1 |
1 |
||
|
Gpd |
b (1.00) |
b (1.00) |
b (1.00) |
e (1.00) |
f (1.00) |
f (1.00) |
a (0.083) |
a (0.500) |
d (1.00) |
||
|
c (0.917) |
d (0.500) |
||||||||||
|
N |
2 |
2 |
7 |
1 |
2 |
1 |
1 |
1 |
|||
|
Gpi |
c (0.500) |
c (1.00) |
c (1.00) |
c (1.00) |
a (1.00) |
d (1.00) |
b (1.00) |
b (1.00) |
|||
|
d (0.500) |
|||||||||||
|
Hbb N |
22 |
2 |
9 |
1 |
2 |
1 |
1 |
1 |
1 |
||
|
100/140/180 |
0.909 |
0.500 |
|||||||||
|
100/140/174/180 |
0.091 |
||||||||||
|
100/140 |
0.500 |
0.889 |
|||||||||
|
100/128/140 |
0.111 |
||||||||||
|
108/140 |
1.00 |
||||||||||
|
100/148/188 |
1.00 |
||||||||||
|
116/144 |
1.00 |
||||||||||
|
64/90/116/144 |
1.00 |
||||||||||
|
96/116/132 |
1.00 |
||||||||||
|
80/100/148/174 |
1.00 |
||||||||||
|
N |
22 |
2 |
9 |
1 |
2 |
1 |
6 |
1 |
1 |
1 |
|
|
Idh-1 |
a (0.477) |
b (0.750) |
b (1.00) |
c (1.00) |
c (1.00) |
c (1.00) |
b (1.00) |
d (1.00) |
d (1.00) |
b (0.500) |
|
|
b (0.523) |
c (0.250) |
c (0.500) |
|||||||||
|
N |
22 |
2 |
9 |
1 |
2 |
1 |
6 |
1 |
1 |
1 |
|
|
Ldh-1 |
c (1.00) |
c (1.00) |
a (0.056) |
c (1.00) |
d (1.00) |
d (1.00) |
c (1.00) |
b (1.00) |
d (1.00) |
d (1.00) |
|
|
c (0.944) |
|||||||||||
|
N |
22 |
2 |
9 |
1 |
2 |
1 |
6 |
1 |
1 |
1 |
|
|
Ldh-2 |
a (0.977) |
a (1.00) |
a (1.00) |
a (1.00) |
d (1.00) |
d (1.00) |
a (1.00) |
c (1.00) |
b (1.00) |
b (1.00) |
|
|
b (0.023) |
|||||||||||
|
N |
22 |
2 |
9 |
1 |
2 |
1 |
6 |
1 |
1 |
1 |
|
|
Ldr |
a (0.227) |
b (1.00) |
a (0.111) |
b (1.00) |
a (1.00) |
b (1.00) |
a (1.00) |
b (1.00) |
a (1.00) |
b (1.00) |
|
|
b (0.773) |
b (0.889) |
||||||||||
|
N |
22 |
2 |
9 |
1 |
2 |
1 |
6 |
1 |
1 |
1 |
|
|
Mdh-1 |
b (0.886) |
b (1.00) |
b (0.833) |
b (1.00) |
b (1.00) |
b (1.00) |
b (1.00) |
a (1.00) |
b (1.00) |
b (1.00) |
|
|
c (0.114) |
c (0.167) |
||||||||||
|
N |
22 |
2 |
9 |
1 |
2 |
1 |
6 |
1 |
1 |
1 |
|
|
Me |
a (1.00) |
a (1.00) |
a (1.00) |
d (1.00) |
c (0.750) |
b (1.00) |
e (1.00) |
a (1.00) |
c (1.00) |
c (1.00) |
|
|
e (0.250) |
|||||||||||
|
N |
22 |
2 |
8 |
1 |
2 |
1 |
6 |
1 |
1 |
1 |
|
|
Np |
b (0.682) |
d (1.00) |
b (0.438) |
a (0.500) |
f (1.00) |
d (0.500) |
e (1.00) |
e (1.00) |
g (1.00) |
i (1.00) |
|
|
d (0.318) |
d (0.563) |
c (0.500) |
e (0.500) |
||||||||
|
N |
22 |
2 |
8 |
1 |
2 |
1 |
5 |
1 |
1 |
1 |
|
|
Pgm |
a (0.023) |
b (1.00) |
b (1.00) |
b (1.00) |
b (1.00) |
b (1.00) |
c (1.00) |
b (1.00) |
b (1.00) |
b (1.00) |
|
|
b (0.977) |
|||||||||||
|
N |
22 |
2 |
9 |
1 |
2 |
1 |
6 |
1 |
1 |
1 |
|
|
Sdh |
a (0.091) |
c (1.00) |
a (0.278) |
f (1.00) |
e (1.00) |
c (1.00) |
b (1.00) |
d (1.00) |
g (1.00) |
g (1.00) |
|
|
c (0.909) |
c (0.722) |
||||||||||
|
N |
21 |
2 |
9 |
1 |
2 |
1 |
6 |
1 |
1 |
1 |
|
|
6Pgd |
a (0.119) |
c (1.00) |
c (1.00) |
d (1.00) |
j (0.750) |
f (0.500) |
g (1.00) |
b (1.00) |
e (1.00) |
e (0.500) |
|
|
c (0.881) |
k (0.250) |
h (0.500) |
g (0.500) |
||||||||
|
N |
22 |
2 |
9 |
1 |
2 |
1 |
6 |
1 |
1 |
1 |
|
|
Sod-1 |
c (1.00) |
c (1.00) |
c (1.00) |
c (1.00) |
b (1.00) |
a (0.500) |
b (1.00) |
b (1.00) |
b (1.00) |
b (1.00) |
|
|
b (0.500) |
|||||||||||
|
N |
22 |
2 |
9 |
1 |
2 |
1 |
6 |
1 |
1 |
1 |
|
|
Sod-2 |
a (1.00) |
a (1.00) |
a (1.00) |
a (1.00) |
a (1.00) |
a (1.00) |
b (1.00) |
a (1.00) |
b (1.00) |
b (1.00) |
N - количество исследованных экземпляров.
Таблица 2 Матрица дистанций Неи (над диагональю) и Роджерса (под диагональю) между таксонами крыс Южного Вьетнама
|
Таксоны |
1 |
2 |
3 |
4 |
5 |
6 |
7 |
8 |
9 |
10 |
||
|
1 |
N.tenaster |
-- |
(0,262) 0,163 |
(0,275) 0,132 |
(0,563) 0,395 |
1,122 |
1,135 |
0,782 |
0,710 |
1,358 |
1,149 |
|
|
2 |
N.fulvescens(1) |
(0,456) 0,375 |
-- |
(0,028) 0,089 |
(0,616) 0,387 |
1,141 |
0,866 |
0,945 |
0,846 |
1,436 |
1,087 |
|
|
3 |
N.fulvescens(2) |
(0,462) 0,334 |
(0,162) 0,284 |
-- |
(0,633) 0,406 |
1,167 |
0,968 |
0,890 |
0,828 |
1,416 |
1,105 |
|
|
4 |
N.langbianis |
(0,630) 0,551 |
(0,665) 0,559 |
(0,668) 0,561 |
-- |
1,004 |
0,980 |
1,066 |
0,996 |
1,432 |
1,083 |
|
|
5 |
M.surifer |
0,788 |
0,811 |
0,802 |
0,781 |
-- |
0,513 |
1,073 |
1,171 |
0,763 |
0,799 |
|
|
6 |
M.moi |
0,781 |
0,740 |
0,752 |
0,767 |
0,612 |
-- |
1,671 |
1,811 |
1,082 |
0,811 |
|
|
7 |
B.bowersi |
0,709 |
0,771 |
0,744 |
0,796 |
0,795 |
0,873 |
-- |
1,135 |
0,869 |
0,839 |
|
|
8 |
L.cf.edwardsi |
0,681 |
0,740 |
0,723 |
0,776 |
0,809 |
0,717 |
0,804 |
-- |
1,011 |
0,981 |
|
|
9 |
R.koratensis |
0,837 |
0,868 |
0,851 |
0,866 |
0,723 |
0,795 |
0,755 |
0,786 |
-- |
0,200 |
|
|
10 |
R.exulans |
0,790 |
0,798 |
0,787 |
0,795 |
0,723 |
0,717 |
0,736 |
0,767 |
0,420 |
-- |
Примечание. Без скобок указаны дистанции, вычисленные по 18 локусам, в скобках -- вычисленные по 27 локусам (см. раздел "Материал и методика")